The heel does not protrude when running.
Humans have to consciously learn a proper technique for running to avoid misusing the heel. Doing "what comes naturally" is just horrible.
Children, when running, do not land on their heels. Heel-landing is a learned development caused by heel-heavy shoes. Children land on the front/middle of their feet.
By contrast, an animal doesn't have to study running technique. A cheetah doesn't have to think "Gee, I'm doing this wrong according to some pages I Googled; I should change my form so that my center of gravity is rather such and such, and my footstrike is altered in such and such a way."
Cheetahs do have to think about how to run. Have you ever seen cheetah cubs?
I should change my form so that my center of gravity is rather such and such, and my footstrike is altered in such and such a way."
Try running without shoes. You'll learn very quickly, without google, not to land on your heels. It's the same way that animals like cheetahs learn to run-by doing.
The sheer thickness of the meat and fat on the lower leg of a human tells you that we have evolved away from running. Running requires a stick-like lower appendage, with all the muscle higher up.
Every claim in that paragraph is wrong.