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Note that this paper is from Nature: Scientific Reports, which is also published by Nature Publishing Group, but is not the Nature journal you're thinking of. SRep is not in the same tier as Nature.

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Thanks for pointing that out.

I'm guessing that it's still a long way removed from "spam", though, right? :-)

Well, SR did have a bit of a credibility issue when they started, and even today, it's not exactly a top journal. I understand it's not a criterion that the articles published are groundbreaking or similar, just that they're decent science and that the authors are willing to pay the hefty fee.

Edit:

See here for one particular discussion, from last year

http://blogs.sciencemag.org/pipeline/archives/2016/06/15/mor...

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