Most of California's agriculture is the Central Valley. Surely that was never desert? It's got rain all winter and runoff from the mountains all summer.
Central Valley farms and the coastal cities are mostly fed by water from the Sierra's, via either rivers or one of California's many aquifers (e.g. Hetch Hetchy, Los Angeles, Colorado, California).
[1]http://en.wikipedia.org/wiki/Central_Valley_(California)#Cli...