>Furthermore, Lightning Network could provide a similar feeless system (excluding the settlement TXs).

There are even ways to get around the extreme majority of the fees for settlement transactions! Look up Channel factories for an idea on how even the opening and settlement transactions can be reduced to about 10% of their current size/cost for a single user. And there might be ways of interacting with channel factories to keep those settlement transactions off the main blockchain for the most part!

Also, I'm fairly certain IOTA doesn't have any TX fees, as well as several others that came before RaiBlocks.