Until you use a non English keyboard.
Alt-F4 is perfectly acceptable OSX has its own share of stupid key bindings especially where the browser is concerned.
I believe the comment about NeXTSTEP was not that the keybindings originated there, but that NeXTSTEP implemented Emacs keybindings in textareas, and that macOS inherited that (I don't know about the accuracy of that though).
Oh, and the page said "Rob Pike answers in 9fans: “I believe ^H ^W ^U date back at least to TENEX.”"; it seems odd to me that he didn't mention that 'H' with the control bit flipped is literally the ASCII "backspace" control character.