I like the question posed at the end of the article: should humans interpret this as a natural expansion or an invasive species. It's hard to tell, that's for sure.
well, for starters, how would humans interpret themselves?
>They thrived, in part, because of increasing forest fragmentation and agricultural lands along with the annihilation of predators like wolves, cougars and jaguars.
sounds like an invasive species [of humans] cleared path for natural expansion of coyotes.