Article is using "memes" interchangeably with "image macros" for those pedants out there like me.
Another criticism comes from semiotics, (e.g., Deacon,[16] Kull[17]) stating that the concept of meme is a primitivized concept of Sign. Meme is thus described in memetics as a sign without its triadic nature. In other words, meme is a degenerate sign, which includes only its ability of being copied. Accordingly, in the broadest sense, the objects of copying are memes, whereas the objects of translation and interpretation are signs.
You can go very far down the rabbit hole of pedantry. ;-)
Edit: spelling