Look at the IE5 screenshot. The second last line has the word "finibus" and notice how the "f" and the "i" seem to be combined together. That's a ligature. In contrast in the body text of the website I don't see any. The word "file" in the paragraph right above the screenshot is missing that ligature.
Kerning is, well, also wrong. So wrong that I literally trouble reading the body text, but I have no trouble reading the text in the IE5 screenshot. (Bad kerning: once you see it you can't unsee it.)