>Analyses of their genomes showed that they contributed genetically to present-day people outside sub-Saharan Africa
This is wrong. Recent analyses (https://www.cell.com/cell/fulltext/S0092-8674(20)30059-3) have shown Neanderthals have also contributed to Sub-Saharan DNA. Maybe the authors weren't aware of the latest findings at the time of writing and no one ever thought of correcting that part.