Lawsuits against Apple's App Store policy enforcement are already in progress, but Apple has the best lawyers money can buy, and rulings are years away.
If a substantial part of your business risked being terminated at the click of a button by some unelected person that you have never met, with no proper way to appeal such an unjust decision, you'd probably think twice about doing something that could be misconstrued as intentionally violating App Store Guidelines too.
I blame Apple for the lack of clarity both in their rules and their enforcement. I don't blame ProtonMail for trying to adhere to Apple's arbitrary rules that often make little sense.