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The Wolves are representative of the Romans!

Legend has it that the founders of Rome were raised by a she wolf! A lot of roman cultural items have an imagery of the she wolf.

https://en.wikipedia.org/wiki/Romulus_and_Remus

Oh, of course. Sigh. Thanks.

The wolves in this case are the Romans themselves (The wolf being a symbol of Rome thanks to the Romulus and Remus myth). republican light infantry wore wolf pelts, and legionary standard bearers wore animal pelt headdresses, including wolves.

Thanks!

And the elephants were explicitly linked to three war-elephant using states absorbed by the Roman Empire, Carthage, Ptolemaic Egypt and the Seleucid Empire.