TL;DR: Combination of brushing scams and forgotten, delayed seed orders.
Here are a few more snippets for those too busy or disinclined to read the whole thing:
"It was now clear to me that at least some of these packets were definitely forgotten orders. ... We couldn’t always come to a definitive conclusion. ... the USDA clearly remained unconvinced by my arguments. ... I certainly don’t rule out there being messier aspects to what happened last summer that remain to be uncovered."