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My dogs growl and yip when wrestling with each other, I feel like this probably comes under the “non-aggressive vocalisation” mentioned in the article.

Growling is obviously a well known sign of fear and aggression but I guess the context is important.

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When playing dogs will indeed bark, growl and whines, but also produce a forced exhalation: that's their laugh. Other sounds can be produced in other contexts indeed, whereas this dog laughter is exclusive to play.

Do you mean the "sorta sneeze" they do? I think mine do that when they think/want that we are going on a walk or into the garden to run about. It's a signal of "let's go adventuring".

There's also the high pitched excited yip and then the tongue out wiggling walk thing as well both of which might be laughter I think.

I read about the “sorta sneeze” as being a call for consensus for “let’s go”. When my dog did that as we were getting ready to go out, I tried making a similar sound back and the dog looked at me immediately like I finally said something she understood!

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