Innovation is born from constraint. Entrepreneurs are relentlessly resourceful because they have to be - not because they can be.
When a VC firm runs on a minimal team, its because it is the preferred model out of many choices. When a startup runs on minimal resources, its because they have no other choice.
The point of lean startup is to focus on learning about customers, and use every conceivable resource to evolve the product team's understanding of how to deliver a great user experience. Things change when the organization becomes well resourced - its not about saving up your pennies anymore to figure out how to learn - its about allocating the resources you have most efficiently to achieve a goal (Occam's razor).
Fred writes awesome posts, but being lean out of necessity will always create a different culture than being lean as a choice and the article did not make this very meaningful distinction.