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People have made harmoniums with modifications to play the 22 shrutis of carnatic music, and with those you can probably more or less approximate the right tones for most hindustani raags too, but it's rare, your average harmonium probably just does bog standard western-style equal temperament.

As the article says though, the main limitation of this kind of instrument is that it cannot generate glissandos (meend) and ornamented notes (gamaka).

They are not difficult to tune and they do tune them but they are used like any other fixed pitch instrument in Indian music, the more easily tuned instruments tune to the more difficult to tune instrument.

Why wouldn't they be? They still use the same 12 notes that correspond to keys on a harmonium.

I’m looking for a different link but this explains it I think

https://chandrakantha.com/forums/?p=post%2Fnotes-on-the-harm...