This is the key problem. To me it looks like most of this article is conflicted about what the definition exactly is:
- basic conscious experiences such as pleasure and pain might not require much species-level intelligence at all
- [there will come] a point at which they also become aware—at which the inner lights come on for them
Based on the text, I don't think I understand what the author believes being concious is. I also don't think this argument will advance further until we start defining precisely what we mean.