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Not in front of my computer right now but I’m pretty sure it’s thumb and index finger. Feels completely natural.

Maybe it has to do with hand size, finger/joint length, etc?

Okay. Otherwise do you in your natural state hold your left hand index on F and right hand index on J? Going to A on pinky on your left hand and the key after L on your right pinky?

Because I think I have pretty much average male hands, but if I have to do CMD and C, with this natural state position, I have to hit the CMD with pretty much the nail of my thumb sideways, unless I rotate my wrist a lot.

on a mac when i'm in mouse and keyboard mode my left thumb is resting on CMD with my index finger on W (CMD+W, close window), middle on 1 (it'll reach down to do CMD+Q), and ` (CMD+`, cycle through windows)

when i'm in programming mode, i'm in emacs on the homerow

Index on F/J, pinky on A/L, thumb naturally resting on space. Hitting left Command involves bending my thumb and though the press is performed with the side of my thumb, it’s not on its nail but further down.

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