>both phenomena are more parsimoniously treated as emergent manifestations of individual phenotypes, and it is individual phenotypes that are directly influenced by genes. You may choose to treat a dominance hierarchy as a group phenotype if you insist, but it is better seen as emerging from each hen, say, being genetically programmed to learn which other hens she can beat in a fight and which normally beat her.

This sounds like an argument that applies equally well to the phenotypes of organisms - you can treat them as entities in their own right, but they're more truly seen as the result of particular gene expressions. Indeed, the end of the article seems to be acknowledging that families have phenotypes; there's no reason this shouldn't apply to groups.