One question I have that I didn't see addressed in the article: how does the inner ear work in low gravity? I wonder if this might make astronauts unusually dizzy on the moon because of differences in the vestibular system.
This article seems like a good survey of the general impact low-g has on the inner ear changes, but I've barely skimmed it: https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8595211/