Honestly, the paper is pretty confusing. It's a weird mix of explaining things people reading a hash paper should already know (e.g. avalanche effect) and not explaining the important bits. The mixing stage looks a lot like a Davies-meyer hash, but it's hard to be sure without reading code. If so, this will be susceptible to fixed point attacks (among others). It doesn't particularly matter because this isn't presented as secure, but worth noting.
The paper also doesn't exhaustively verify numbers when domain=codomain, which is a super common scenario and a weird omission. 32 bits is small enough to check everything.