Yes, saving 1% of a few million requests a day is not a trivial savings.
But I don't think we need to think of it in terms of Twitter's scale at all. The point is that saving 1% for basically free, is saving 1% for basically free, and that's worthwhile no matter how big you are. On its own, it's a drop in the bucket, but the cumulative effect of many small low-opportunity-cost savings is a significant low-opportunity-cost saving.