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If developers and enterprises can host their own OSS/fine-tuned models, why will they pay Anthropic or OpenAI?

Because hosting a really GOOD model requires hardware that costs tens of thousands of dollars.

It's much cheaper to timeshare that hardware with other users than to buy and run it yourself.

That may be true for independent devs or startups. Larger companies have enough demand to justify a few A/H100s.