I would assume that his suggestions for clarity in "scientific papers" and his literary style don't overlap all that much to infer the former from the later.
> Commas denote a pause in speaking.... Speak the sentence aloud to find pauses.
made its way into this article. Hard to imagine that this particular point, to which I might attribute many of the comma splices I see in scientific writing, actually came from a professional writer.
my writing advice:
never use the former and the latter