I described what (I think) I'm seeing in https://github.com/Rabrg/artificial-life/pull/1 , and suggested in https://github.com/Rabrg/artificial-life/pull/3 that random mutations ought to blow away the whole tape rather than modify only one cell of it. However, even with that modification to the rules, I still see the `<` substrate taking over the whole map, except when the mutation rate is very high. (I admit this doesn't make a ton of sense to me.)
So, anyway, it's easy to make the mutation rate vary from one side of the map to the other. If you do that right, you can see the `<` substrate take over the low-mutation-rate side of the map, but fail to take over the higher-mutation-rate side.
The `<` substrate is easy to see in the gif: it's a rapidly spreading splotch of red with gold noise flickering inside.