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Does that mean soon I'll be able to control playback speed on Coursera and Khan Academy (presuming I've signed up for HTML5 beta in YouTube, which I have)? Anyone know?

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If this new H.264 support in Firefox means that you can control playback speed on YouTube itself, you should also be able to control the speed on Khan Academy (or any other site that uses YouTube's iframe embed).

Support for HTML's playbackRate API was landed a few weeks back. See bug 495040 https://bugzilla.mozilla.org/show_bug.cgi?id=495040

This means that sites that control playback speed using the HTML media API should work.

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