I am an engineer at Tokutek. Just talking about TokuMX, I have only seen MongoDB beat us on single-threaded, read-only workloads on extremely small data sets (like under 8K, or one MongoDB B-tree bucket).
The statement is really pretty much true. The best B-tree indexes (InnoDB) beat Fractal Tree indexes a little bit on in-memory reads, but we're not done tuning our implementation. On out-of-memory reads though, we usually beat B-trees because we compress so much better that we simply need to read less off disk.
Benchmarking is a tricky business though. Workloads can be varied, and you can probably find some corner cases where InnoDB beats TokuDB. I think we have some outstanding issues where the MySQL optimizer doesn't plan our queries properly, and we're still working on that. I'm a little out of touch with the MySQL side these days though so that could be wrong. But algorithmically, there are no cases where a B-tree index has a significant advantage over a Fractal Tree index.
Generally though, the read performance advantage we see is that if your indexes are Fractal Tree indexes, you can afford to maintain a richer set of indexes than you could have on InnoDB or MongoDB, and these extra indexes make your queries orders of magnitude faster. I think this is the most important (non-obvious) point to understand. I gave a talk about it here: http://www.youtube.com/watch?v=q6BnG74FZMQ