A decent number of people at the top of the food chain in Silicon Valley come from finance, a world very much driven by pedigree. We tend to forget that from time to time, most of us here (I would imagine?) being engineers, and fewer of us being financiers. But pedigree has always counted in finance. Though it counts
slightly less today than it did 30 years ago, it still counts for quite a lot.
To this day, there are circles one simply can't break into unless one comes from a top Ivy/Stanford undergraduate program or business school. These circles dominate recruiting at top investment banks and consultancies, which in turn feed VC firms and executive positions at many tech companies. It's not impossible to land one of these roles without a degree from, say, Harvard Business School. But that degree will open doors that are, by default, closed to most everyone else without extraordinary hustle.
Another very important point: many of those doors opened on campus, not afterward. Having Harvard, Yale, Stanford, etc., on your resume looks lovely. But that's almost incidental to the real benefit. The real benefit is that the top banks, consulting firms, and recruiters show up on those campuses and pay greater attention, hire more interns, and so on. The advantages begin at school and start accumulating afterward. This is the secret sauce that MOOCs will have a hard time replicating, at least until the top firms become more open-minded about recruiting.
The importance of pedigree will change when the demand for pedigree changes, not the supply. This is why I remain optimistic about the prospects of online courses in the long run, but skeptical of their prospects to really shake things up in the short run. So long as top-tier job recruiting demands pedigree, pedigree will matter.
I think demand-side investment by organizations like YC, 500 Startups, etc. -- many of which seem fairly indifferent to pedigree -- will make a bigger impact than will supply-side attempts such as MOOCs or their equivalents. At least for now.