other option is to have a Printer(Human)-Friendly web page button, don't know if reviewers have this option, if isn't available i think a Human-Friendly layout could reduce fatigue from reading so many applications.
I can only speak for myself, this would be a feature i would like to have.
I would have loved one: it's easier to share the application with co-founders and have them annotate it, before editing.
One solution is to write the YC application using your favorite word processor (vi,Notepad,Word), and when finished copy paste the content to the application form.