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I think YOOTheme's UIKit is a great modular framework that not many people know about.

Agreed. I was surprised to not see it mentioned in the article, and only at the bottom here. I love it, it's my goto front-end framework.

What are the advantages of UIKit over bootstrap?

It's small, without over-complicated class names and has all kinds of add-ons like dropdown etc. available as modules.

And also – easily tweakable.