i don't propose a "main culprit" as i understand climate to be a classic complex system. i simply don't know what the main culprit is or even if there IS a main culprit.
i'm not yet convinced that the temperature increase that we've seen in the past 150 - 200 years is abnormal from a geologic timeframe. esp. considering the warming pause of the past ~20 years.
as far as the article you posted, i've read it before. it's fairly standard in that it adds another theory (southern warming -> release co2 -> increase northern warming) that, for me, would require more observable predictions to prove. that's not an easy task, but my scientific skepticism requires it.
maybe if we could see that effect happening in our modern temperature data? i'm just riffing here, but it seems like if that's the mechanism to explain the ~800 year lag where temperature increase preceeds co2 increase, then it should also be observable today.