The trouble I have with much of this discussion is it assumes many traits of animal behavior are indicative of consciousness. Bees play with balls, therefore consciousness? You can't just jump straight to that conclusion. There is so much our brains do that is subconscious, that we are not aware of. Heck, even signs of self awareness may not be indicative of consciousness. Arguably we are at the point that we could train a robot AI to recognize itself in the mirror.
I'm in the camp that believes animals are conscious, I'm not arguing against that. But what I'm arguing is that the current body of evidence doesn't conclude anything. Until we have a better fundamental understanding of consciousness, I don't think we can make such conclusions merely from observing animal behavior. Who's to say that one behavior is evidence of consciousness and another behavior is not? What do we base that upon?