The article says the spheres will be at 500-600m depth, so something like 50-60 atmospheres of pressure. Compressing air enough to displace water at that pressure would heat it to extreme temperatures which must be very inefficient even with some attempt to recover the heat. Water being incompressible can I assume be moved in and out of the sphere without any heating (and, therefore, energy loss).
(I'm guessing, of course)