I encoded the gif in 420 (subsampling), 422 and 444, having a nice quality with 444 with VLC, I embed the three different videos in a web page and when playing in my browser, they are all as bad as the twitter video! I thought that Firefox was using a standard library to play MP4?
Depending on the webpage you uploaded them, the videos could be re-encoded (or VLC's options could be no-ops and the artifacts hidden when played back in VLC due to better chroma upsampling) hence the problem.
When I said that I embedded the videos in a web page, I wrote a small page from scratch to display them in my browser, locally, without any server. So no re-encoding. I think that VLC just has a better chroma upsampling.