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Firefox uses the Windows-bundled decoder on Windows Vista or later. Chroma upsampling happens after decoding so it's the renderer's responsibility (Firefox, Chrome, VLC, madVR etc).

Depending on the webpage you uploaded them, the videos could be re-encoded (or VLC's options could be no-ops and the artifacts hidden when played back in VLC due to better chroma upsampling) hence the problem.

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Okay, thanks for the explanation on the chroma upsampling! I use Linux but it should work all the same.

When I said that I embedded the videos in a web page, I wrote a small page from scratch to display them in my browser, locally, without any server. So no re-encoding. I think that VLC just has a better chroma upsampling.

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